I'm making an element using spikeviper's biology mod, and it's resulting in a strange error I can't find helpful info about.

Source code:

#include "simulation/ElementCommon.h"

static int update(UPDATE_FUNC_ARGS);
static int graphics(GRAPHICS_FUNC_ARGS);

void Element::Element_TUMOR()
{
    Identifier = "DEFAULT_PT_TUMOR";
    Name = "TUMR";
    Colour = PIXPACK(0x770000);
    MenuVisible = 1;
    MenuSection = SC_BIO;
    Enabled = 1;

    Advection = 0.0f;
    AirDrag = 0.00f * CFDS;
    AirLoss = 0.97f;
    Loss = 0.50f;
    Collision = 0.0f;
    Gravity = 0.0f;
    Diffusion = 0.00f;
    HotAir = 0.000f    * CFDS;
    Falldown = 0;

    Flammable = 0;
    Explosive = 0;
    Meltable = 0;
    Hardness = 12;

    Weight = 150;

    DefaultProperties.bio.health = 150; // In real life, cancer survives better than regular tissue (citation: HeLa)
    DefaultProperties.temp = R_TEMP - 2.0f + 273.15f;
    HeatConduct = 29;
    Description = "Tumor. Caused by bombarding biology with neutrons";

    Properties = TYPE_SOLID|PROP_NEUTPENETRATE|TYPE_BIO;

    LowPressure = IPL;
    LowPressureTransition = NT;
    HighPressure = IPH;
    HighPressureTransition = NT;
    LowTemperature = ITL;
    LowTemperatureTransition = NT;
    HighTemperature = ITH;
    HighTemperatureTransition = NT;

    Update = &update;
    Graphics = &graphics;
}

static int update(UPDATE_FUNC_ARGS)
{
    int r, rx, ry;

    rx =  RNG::Ref().between(-2, 2);
    ry =  RNG::Ref().between(-2, 2);

    // O2 use by tumor itself (Increased due to increased biological activity)
    if (RNG::Ref().chance(1, 100)){

        if (parts[i].bio.o2 > 0){
            parts[i].bio.o2 -= 1;
                parts[i].bio.co2 += 1;
        }
    }


    if (BOUNDS_CHECK && (rx || ry))
    {
        r = pmap[y+ry][x+rx];
        if (r) {
            if (RNG::Ref().chance(1, 2)){
                // Diffuse among tumor
                if (TYP(r) == PT_TUMOR){
                    int ir = ID(r);

                    if (parts[i].bio.o2 > parts[ir].bio.o2){
                        parts[i].bio.o2--;
                        parts[ir].bio.o2++;
                    }
                    if (parts[i].bio.co2 > parts[ir].bio.co2){
                        parts[i].bio.co2--;
                        parts[ir].bio.co2++;
                    }
                }
                // steal o2 from bio, offload co2 to bio (tumor is greedy)
                if (sim->elements[TYP(r)].Properties & TYPE_BIO){
                    int ir = ID(r);

                    if (parts[ir].bio.o2 > 1){
                        parts[i].bio.o2++;
                        parts[ir].bio.o2--;
                    }
                    if (parts[i].bio.co2 > 0){
                        parts[i].bio.co2--;
                        parts[ir].bio.co2++;
                    }
                }
                if (RNG::Ref().chance(1, 50)){
                    // convert biology to tumor (grow)
                    if (sim->elements[TYP(r)].Properties & TYPE_BIO && TYP(r) != PT_TUMOR){
                        int ir = ID(r);
                        sim->part_change_type(ir, parts[ir].x, parts[ir].y, PT_TUMOR);
                    }
                }
            }
        }
    }

    // Health management
    if (RNG::Ref().chance(1, 50)){
        // Temp check
        if (parts[i].temp > 323.15){
            int damage = (parts[i].temp - 315) / 5;
            parts[i].bio.health -= damage;
        }
        // Damage check
        if (parts[i].bio.co2 > MAX_CO2 || parts[i].bio.o2 < 1){
            parts[i].bio.health--;
        }
        // Otherwise heal
        else{
            // Tumors aren't the most healthy to start with.
                        if (parts[i].bio.health < 200){ 
                parts[i].bio.health++;
            }
        }
    }

    // Death check
    if (parts[i].bio.health < 1){
        sim->part_change_type(i, x, y, PT_DT);
    }

    return 0;
}

static int graphics(GRAPHICS_FUNC_ARGS)
{
    // Oxygen
    int o = cpart->bio.o2;

    // C02
    int c = cpart->bio.co2;

    *colr = (int)fmax(7 * o, 100);
    *colg = 0;
    *colb = (int)fmax(3 * o, 30);
    *pixel_mode |= PMODE_BLUR;

    // Life mix
    *colr = int(*colr * (cpart->bio.health) / 100.0f);
    *colg = int(*colg * (cpart->bio.health) / 100.0f);
    *colb = int(*colb * (cpart->bio.health) / 100.0f);

    return 0;
}

and at https://github.com/SpikeViper-Powdertoy-fork/The-Powder-Toy/blob/master/src/simulation/elements/TUMOR.cpp

The error is

../src/simulation/elements/TUMOR.cpp:107:4: error: expected primary-expression before ‘}’ token

and I couldn't find any relevant info, except for a thing telling me to check the curly braces, which I already did. So, can someone explain how to fix it? (107 is at the middle of the big stack of five closing curly braces)

Premptive Thank You!